More than 300 tiny pieces of human tissue from prisoners executed in Nazi-era Berlin will be buried on Monday.
The samples were found in microscopic slides at a property that belonged to Hermann Stieve – an anatomy professor at the Charité university hospital.
Heirs of the doctor, who died in 1952, discovered the collection in 2016.
Researchers say Stieve systematically collaborated with the Nazis to receive the bodies of 184 people, mostly women, executed for political resistance.
The tissue pieces – most less than a millimetre long – were discovered at Stieve’s estate, stored in small black boxes, including some labelled with names.
Once found, they were handed to Berlin’s Charité university hospital, who tasked staff at the German Resistance Memorial Center to research their history.
Research under the memorial’s director, Prof Johannes Tuchel, showed that bodies were picked up by a driver and taken to Stieve, sometimes just minutes after they were killed at Berlin-Plötzensee prison.
He then dissected them for research, before discreetly cremating and interring their bodies anonymously.
Almost 3,000 people were executed at Plötzensee by beheading or hanging while Hitler was in power.
“We have discovered that (Stieve) systematically aided the (Nazi) Reich Justice Ministry in obliterating the traces of these criminal acts,” Prof Tuchel told German newspaper Bild.
Stieve served as the director of the Berlin Institute of Anatomy from 1935 until he died following a stroke in 1952.
The anatomist’s use of the prisoners’ corpses had been kept almost in plain sight, because he kept meticulous records of his work.
He had a particular interest in reproductive anatomy.
His work was some of the first research to suggest that stress – in the form of being sentenced to death – could disrupt a woman’s menstrual cycle.
Some of the people he dissected were high-profile – including 13 women from the Red Orchestra communist resistance group.
One of the project’s researchers was Andreas Winkelmann, a department head at the Institute of Anatomy of the Brandenburg Medical School (MHB).
He told the AFP news agency that burial of such small specimens was highly unusual.
“But this is a special story, because they came from people who were actively denied graves, so that their relatives would not know where they are buried,” he added.
Dr Sabine Hildebrandt is a German-born anatomist who published a book about ethical transgressions and anatomical science in the Nazi period.
In 2013 she explained to the BBC that Stieve exploited their policies, including the increased use of the death penalty as a punishment.
“Before 1933, he was able to source the bodies of executed men, but no women; Germany was not executing women,” she said.
“Then, suddenly, during the Third Reich, women were being executed too.”
Because he was not a member of the Nazi party, Stieve was not prosecuted after World War Two.
The samples will be laid to rest at a ceremony at Berlin’s Dorotheenstadt Cemetery on Monday.
In a statement, Dr Karl Max Einhäupl, CEO of the Charité, said the burial was part of an effort by the hospital to confront its – and German medicine’s – difficult relationship with Nazism.
“By burying the microscopic specimens at the Dorotheenstadt Cemetery, we want to help restore to the victims some of their dignity,” he said.